Anatomy And Physiology Of The Female Reproduction System.

 

Female reproductive systems

 

 



 

Female reproductive systems consist of the following

 

1)   External genitalia

2)   A pair of primary sex glands (gonads)

3)   Ducts leading from the gonads to the body’s exterior

4)   secondary sex glands.

In the normal course of events, life begins and is sustained for 9 months within the productive environment of the female. Therefore the female organs will

 

 

 

be considered first, beginning with the external genital structures.

 

The female’s external and internal reproductive structure are targets for estrogen’s throughout her life span and they atrophy (reduce in size) with age and /or a drop in estrogen production.  An extensive and complex innervation and very generous blood supply support the functions of these structures.  The comparative size, woman depending on her heredity, age, race and number of children she has borne.

External Structures

 

Following are the external female genitalia (vulva, pudenda).

 

1)   Mons veneris (mons pubis)

2)   Labia majora and minora

3)   Clitoris

4)   Vestibule

¨1  Urinary meatus, or urethral orifice

¨2  Lesser vestibular, paraurethral, or Skene’s glands

¨3  Hymen and vaginal introitus, or orifice

¨4  Greater vestibular, vulvovaginal, or Bartholin’s glands

1)   Fourchet

2)   Perineum

 

The mons veneris or mons pubis, is the rounded, soft fullness of subcutaneous fatty tissue and loose connective tissue over the symphysis pubis. It contains many sebaceous (oil) glands and develops coarse, dark, curly hair at pubarche, about 1 to 2 years before the onset of the menses.

 

Typically, the pattern of hair growth in 75% of women is a triangle shape with the base along the top of the symphysis pubis. In 25% of women, pubic hair extends upward the umbilicus along the linea alba. The function of the mons are to play a role in sensuality and to protect the symphysis pubis during coitus.

 

Labia majora are two rounded lengthwise folds of skin-covered fat and connective tissue that merge with the mons. They extend from the mons downward around the labia minora, ending in the perineum in the midline, and act as protection for the labia minora, urinary meatus, and vaginal introitus.

 

On their lateral surfaces, the labial skin is thick, usually pigmented darker than the surrounding tissue, and covered with coarse hair that thins out toward the perineum. The medial (inner) surfaces of the labia majora are smooth, thick, and without hair. They contain an abundant supply of sebaceous glands and sweet glands and are highly vascular.

The lymphatic system is extensive and serves many structures in the pelvic area. This is the reason for the extensive and rapid spread of reproductive tract malignancies.

 

The extreme sensitivity of the labia majora to touch, pain and temperature is due to the extensive network of nerves. Innervation of the anterior one third of the labia is from L1 (lumber 1), while the posterior two thirds is supplied mainly from S3 (sacral 3)

 

The pair of round ligaments from the uterus inserts into the anterior portion of the labia majora.

The clitoris is a short, cylindric, erectile organ liked just beneath the arch of the pubis; the visible portion is about 6 X 6 mm.

Its rich vascularity and innervation make the clitoris highly sensitive to temperature, touch, and pressure sensation.

 

The vestibule is an ovoid or boat-shaped area formed between the labia minora, clitoris, and fourchet.  The vestibule contains the openings to the urethra, paraurethral (lesser vestibule, Skene’s ) glands, the vagina, (greater vestibular or Bartholin’s) glands.

 

The humen is a partial, rarely complete, elastic but mucosa-covered fold around the vaginal  introitus

 

The greater vestibular (Bartholin’s) glands are two compound glands at the base of the labia majora, one on either side of the vaginal orifice. Each gland is drained by several ducts, about 1.5 cm long. Each opens into the groove between the hymen and the labia minora. Usually the gland openings are not visible or palpable. The glands secrete a small amount of clear, viscid mucus, especially during coitus. The alkaline pH of the mucus is supportive of sperm. These glands are susceptible to gonorrheal infection and to abscess and cyst formation.

 

The fourchet is a thin, flat, transverse fold of tissue formed where the tapering labia majora and minora merge in the midline below the vaginal orifice.

The perineum is the skin-covered muscular area between the vaginal introitus and the anus. The perineum forms the base of the perineal body.

 

Pelvic floor and perineum. The pelvic and perineum are composed of the pelvic diaphragm, the urogenital diaphragm or triangle, and the muscle of the external genitalia and anus

 

The upper pelvic diaphragm ,composed of muscles and their fascia and ligaments, extends across the lowest part of the pelvic cavity. The largest anus most significant portion of the diaphragm is formed by the pair of broad, thin levator ani muscles that extend sheetlike between the ischial spines and coccyx, and the sacrum. The levator ani group of muscle is made up of three muscle pairs:

·        puborectalis,

·        iliococcygeus, and

·        pubococcygeus muscles.

The pubococcygeus muscle is particularly significant for women. Its importance lies on bladder control and during labor for controlling perineal relaxation and expulsion of the fetus during birth.

 

The second paired muscles of the upper pelvic diaphragm are the closely joint coccygeus muscles  These muscles extend from the ischial spines to the coccyx and lower sacrum.

 

The urogenital (lower pelvic) is located in the hollow of the pubic arch and consists of the transverse perineal muscles. The transverse  perineal muscles originate at the ischial tuberosities and insert into the perineal body. The strong muscle fibers provide support to the anal canal during defecation and to the lower vagina during delivery. The deep transverse perineal muscles join to form a central seam or raphe. Some of their fibers encircle the urinary meatus and vaginal sphincters.

 

The perineum is located below the upper and lower pelvic diaphragm. Its muscles and fascia reinforce the strength of the pelvic diaphragm and aid in constricting the urinary, vaginal, and anal openings. The bulbocavernosus muscle fibers originate in the perineal body and surround the vaginal openings as the muscle fibers pass forward to insert into the pubis.

The ischiocavernous muscle originate in the tuberosities of the ischium and continue at an angle to insert next to the bulbocavernosus muscle. These muscle fibers contract to cause erection of the clitoris.

Anal sphincter muscle fibers originate at the coccyx, separate to pass on either side of the anus, fuse, and then insert into the transverse perineal muscles.

The bulbocavernosus, transverse perineal, and anal sphincter muscle fibers can be strengthened through Kegel exercises.

The perineal body, the wedge-shaped mass between the vaginal and anal openings, serves as an anchor point for the muscle, fascia, and ligaments of the upper and lower pelvic diaphragm The skin-covered base of the body is known as the perineum. The perineal body, about 4 cm wide by 4 cm deep, is continuous with the septum between the rectum and vagina.

Knowledge of the anatomy and physiology of the pelvic floor and perineum provides a basis for understanding and implementing appropriate nursing actions in relations to voluntray sphincter control during urination, coitus, and defecation and the normal mechanism of labor and delivery. In addition, the student has the information necessary to understand and utilize knowledge regarding the etiology, management, and prevention of disorders such as uterine prolapse and urinary stress incontinence.

 

Internal Genitalia

 

Vagina

 

Location and support. The vagina is a tubular structure located in front of the rectum and behind the bladder and urethra. The vagina extends from the interoitus, the external opening in the vestibule between the labia minora of the vulva, to the cervix. It is supported mainly by its attachment to the pelvic floor musculature and fascia.

 

Structure. The vagina is a thin-walled, collapsible tube that is capable of great distention. Because of the way the cervix protrudes into the uppermost portion of the vagina, the length of the anterior wall of the vagina is only about 7 to 8 cm while that of the posterior wall is about 10 cm. The recesses formed all around the protruding cervix are called fornices: right, left, anterior, and posterior. The posterior fornix is deeper than the other three.

The smooth muscle walls are lined with glandular mucosa is arranged in transverse folds called rugae.

Innervation. The vagina is relatively insensitive. There is some innervation from the pudendal and hemorrhoidal nerves to the lowest one third. The nerve supply is mainly autonomic. Sensations arising in the vagina terminate at the level of S2, S3, and S4.

Blood supply. The copious blood supply to the vagina is derived from the descending branches of the uterine artery, the middle hemorrhoidal artery, and the internal pudendal arteries. The venous return of the vaginal blood is through the pudendal, external hemorrhoidal, and uterine veins.

Lymphatics. The lymphatics of the upper vagina drain to the rectovaginal septal, presacral, external iliac, and hypogastric nods. The lower vaginal lymphatic are directed to the superficial inguinal nodes.

 

Uterus

The uterus is a hollow viscus composed of plain muscle whose sole function is gestation.  It lies between the rectum and the bladder and is continuous with the vagina.

 

 

Structure

 

Shape, size, and divisions. The uterus is a fattened, hollow, muscular, thick-walled organ that looks somewhat like an upside-down pear. Its length, width, and thickness vary, averaging about 7.5 cm x 3.5 cm x 2cm (3 x 11/2 x 3/4 in). In adult woman who has never been pregnant, the uterus weighs 60 g (2 oz).

The uterus has three parts :the fundus, the upper, rounded prominence above the insertion of the fallopian tubes; the corpus, or main portion, encircling the intra-uterine cavity; and the isthmus, the slightly constricted portion that joins the corpus to the cervix and is known as the lower uterine segment during pregnancy.

Uterine wall. The wall of the uterus is made up of three layers: the endometrium, the myometrium, and a partial outer layer of parietal peritoneum.

Cervix The lowermost portion of the uterus is the cervix, or neck  The attachment site of the uterine cervix to the vaginal vault divides the cervix into the longer superavaginal (above the vagina) portion and the shorter vaginal portion. The length of the cervix is about 2.5 to 3.0 cm, of which about 1 cm protrudes into the vagina in the nongravid woman.

The external os is small before parturition. After the birth of a child it becomes a transverse slit ‘the parous os’

 

Uterine position. For the majority of normal women, with the urinary bladder empty, the uterus is anteverted and slightly anteflexed. The cervix is directed downward and backward the tip of the sacrum so that it is usually at approximately a right angle to the plane of the vagina. For other women the uterus may be in the middle position or tipped backward (retroverted)

A full bladder pushes the uterus back toward the rectum, while a full rectum moves the uterus forward against the bladder. Uterine position also changes depending on the woman’s position (e.g., lying supine, prone, on her side, or standing), her age, and pregnancy.

Uterine support.  The uterus is supported by ligaments and by muscles of the pelvic floor, including the perineal body. A total of 10 ligaments stabilize the uterus within the pelvic cavity : four paired ligaments (broad, round, uterosacral, and cardinal) and two single ligaments (anterior and posterior).

 

The paired broad ligaments are double folds of parietal peritoneum that extend wing-like from the sides of the uterus to the pelvic walls. These ligaments divide the pelvic cavity into anterior and posterior components. In the upper portion of the broad ligaments are suspended the fallopian tubes, ovaries, round ligaments, and ovarian ligaments.

 

The two round ligaments extend from the upper outer angles formed where the fallopian tubes join the uterine corpus (at the cornua), through the inguinal canals, and ending in the labia majora.

The single anterior (uterovesical or pubocervical) ligament is a continuation of parietal peritoneum that forms the anterior fold of the broad ligament, extending from the anterior surface of the supravaginal cervix of the uterus to the posterior surfac4e of the bladder. The pouch formed by the fold of peritoneum is less deep than that the posterior pouch.

The denser connective tissue of the lower portion of the broad ligaments is sometimes known as the cardinal, transverse, or Mackenrodt’s The cardinal ligaments form the upper portion of the posterior ligament.

The single posterior (or rectovaginal ) ligament is a continuation of parietal peritoneum (posterior fold of broad ligament) extending from the posterior surface of the uterus to the rectum. The posterior ligament forms the deep rectouterine pouch also known as the cul-de sac of Douglas

This pouch is the lowest part of the abdominal cavity so that blood, pus, or other drainage collects here. The pouch can be reached through the posterior vaginal fornix.

The two uterosacral ligaments are cord like folds of peritoneum extending from the supravaginal cervical portion of the uterus to the fascia over the second and third sacral vertebrae passing on each side of the rectum. These ligaments hold the uterus in position by maintaining traction on the cervix.

In summary the main uterine support are the ligaments surrounding the supravaginal cervix:

1)    Anterior (pubocervical)

2)   Cardinal (transverse, Mackenrodt’s)

3)    Posterior (rectovaginal)

4)    Uterosacral

 

Uterine lymphatics. The lymphatics of the uterus are extensive. They are contained in three networks: at the base of the endometrium, within the myometrium, and just under the peritoneal coat of the uterus. Lymphatic drainage occurs mainly at the isthmus along the uterine vessels.

Uterine blood supply. The abdominal aorta divides at about the level of the umbilicus and forms the two iliac arteries. Each iliac artery divides to forms the two iliac arteries, the major one of which is the hypogastric artery. The uterine arteries branch off from the hypogastric arteries.

In the nonpregnant state the uterine blood vessels are coiled and tortuous. With advancing pregnancy and an enlarging uterus, these blood vessels straighten out. The uterine veins follow along the arteries and empty into the internal iliac veins.

Innervation  of the uterus. The internal genitalia have a rich supply of afferent and efferent autonomic nerves both motor and sensory.

Motor nerves. Parasympathetic fibers from the sacral nerves are probably responsible for producing vasodilatation and inhibiting muscular contraction. Efferent sympathetic motor nerves arise from the ganglia of T5 (thoracic5) to T10, come together over the sacrum, and reach the uterus through ganglia that lie near the base of the uterosacral ligaments. The autonomic nerves just described (parasympathetic fibers and efferent sympathetic motor) regulate the action of the uterus, but the uterus has an intrinsic motility (i.e., it can contract and relax even if the nerves to it are cut).

Sensory nerves. Sensory fibers carrying pain sensation from the uterus, come together in the paracervical areas and proceed upward to pass just below the division (bifurcation of the aorta, and then travel into the spinal cord at the level of T11 and T12. Because of this arrangement, pain that originate in the ovary or in the ureters may mimic pain that originate in the uterus, any of which may be felt in the flank and down to the inguinal and vulvar areas.

Functions. The three functions of the uterus are essential for the survival of the species but not for the individual: cyclic menstruation with rejuvenation of the endometrium, pregnancy, and labour.

Fallopian tubes. The paired Fallopain (uterine) tubes   measure about 12 cm in length. Each tube has an outer coat of peritoneum, a middle thin muscular coat, and an inner mucosa.  The smooth muscle fibers are arranged in an inner circular and an outer longitudinal layer. The mucosal lining consists of columnar cells some of which are ciliated and others of which are secretory.

The structure of the fallopian tube changes along its length.

Four distinctive segments can be identified:

1.      the infundibulum,

2.      the ampulla,

3.      the isthmus, and

4.      the interstitial part.

The infundibulum, is the most distal portion. Its funnel-shaped opening is encircled with fimbria. The fimbria become swollen, almost erectile, at ovulation. It is 2cm long. The function of these fingerlike projections is to pull the ovum into the tube.

The ampulla makes up the distal and middle segment of the tube. It is in the ampulla that the sperm and ovum meet, where ferilization occurs. 5cm long, thin walled and convoluted.

The isthmus is proximal to the ampulla. It is 2 cm long, straight and cord-like 1mm diameter. 

The interstitial portion passes through the myometrium between the fundus and the body of the uterus and 1cm long and very narrow (less than 1mm).


Ovaries: female gonades

 

 

 


 


Location and support. One ovary is located on each side of the uterus, below and behind the uterine tubes. The ovaries are held in place by two ligaments, the mesovarian portions of the uterine broad ligament, which suspend them from the lateral pelvic side walls at about the level of the anterosuperior iliac crest, and the ovarian ligaments, which anchor them to the uterus. The ovaries are movable with palpation.

 

Structure. Each ovary is composed of two layers around a central zone. Each is whitish and rounded but flattened weights about 3 g, and measures approximately 3 x 2 x 1 cm. At the time of ovulation, ovarian size may double temporarily.

Blood supply and lymphatics. Ovarian arteries carry a rich blood supply from the aorta to the ovaries. The left ovarian vein empties into the left renal vein, but the right drains into the inferior vena cava. The ovarian lymphatics drain into the iliac and periaortic nodes.

Innervation. The nerve supply to the ovary is through T10 to L1, together with fibers of the pelvic sympathetic nervous system.

Functions. The two functions of the ovary are ovulation and hormone production.

Anatomy of the bony pelvis. During birth, the fetus must pass through the bony passage formed by the pelvis

The pelvis is a bony ring that is supported by the lower extremities and that in turn the weight of the trunk and upper body.

The pelvis is made up of the following four bones: the right and left innominate bones, each of which is made up of the right or left pubic bone, ilium, and ischium, which fuse after puberty; the sacrum , and the coccyx.

The two innominate bones (hip bones) form the sides and front of the bony passage, and the sacrum and coccyx form the back.

 

Below the ilium is the ischium, a heavy bone terminating posteriorly in the rounded protuberances known as the ischial tuberosities. The tuberosities bear the body’s weight in the sitting position.

 

The sharp projections, the ischial spines, project from the posterior border of the ischium into the pelvic cavity. The spines, which may be blunt or prominent, have obstetric importance for two reasons:

1)    The distance between the two ischial spines is the narrowest diameter of the pelvic cavity.

2)    They serve as the landmark to determine the degree of descent of the fetus during labor.

The pubis, forming the front portion of the pelvic cavity, is located beneath the mons. In the midline, the two pubic bones are joined by strong ligaments and a thick cartilage to form the joint called the symphysis pubis. The descending rami (ramus [sing.] branch or arm) form the subpubic arch. In the female, the angle formed by the arch optimally measure slightly more than 90 degrees.

The sacrum is formed by five fused vertebrae. The upper anterior portion of the body of the first sacral vertebra, the promontory, forms the posterior margin of the pelvic brim. The

contents of the peritoneum (culdocentesis).

Following the examination, the physician, using diagrams or pictures as indicated, should discuss the findings with the patient so that she understands her condition and the alternatives of therapy.

 

Menstrual Cycle

 

Menstrual Myths

An awareness of some myths about menstruation and their cultural origins is necessary to use the nursing process effectively with both female and male clients.

The belief systems held by researchers may also influence the research methodology used and the way results are interpreted, presented in the literature, and used by society in general (i.e., people tend to color things the way they want to see them).

 

Many myths have their origin in the mystery that surrounded the woman, her hidden reproductive organs, and her uniqueness in adding new members to society. Before the eighteenth century, the relationship of the menstrual cycle to reproduction was obscure.

Menstrual flow was believed to be the elimination of poisons that had accumulated in the woman. The discovery of the fallopian tubes did nothing to clear up this misconception.

 

Because of their recurring nature, menstrual cycles were thought to be under the control of the moon. Before the discovery of ovulation in humans, it was thought that an egg was produced during menstruation only when fruitful intercourse had occurred. Not until the nineteenth century was knowledge available about the existence of the human egg, ovulation and ovarian functioning.

The most common myths in existence today can be sorted out under the following headings:

 

1.      During menstruation, the woman is vulnerable and therefore needs to be protected.

2.      The menstruating woman can pose a danger.

3.      Menstrual blood has been  viewed as possessing healing powers. Or of some black magic power

 

The menstruating woman is seen as being vulnerable to physical and psychological stress. Recall some of myths you may have heard; for example, “Don’t wash your hair,” “Don’t take a bath,” “Watch out, you’ll catch cold,” “That’s too heavy for you to you to carry now ,”

 

Historical Perspective

 

During this century there have been significant additions to the knowledge of the biophysical and emotional aspects of the menstrual cycle. Since then, pituitary hormones have been purified and hypothalamic control over the pituitary gland has been studied extensively.

 

Hypothalamus and Pituitary

 

Neurons from several portions of the brain terminate in the hypothalamus. The cell bodies of the secretory neurons are located within the median eminence (middle portion) of the hypothalamus while the nerve endings lie close to capillary loops slightly above the pituitary portal (circulatory) system. Thus the circulation picks up the hypothalamic neurohormones and carries them to the anterior pituitary, which normally responds to each specific releasing or inhibiting neurohormone “command.”

One releasing hormone, gonadotropin-releasing hormone (GnRH) causes the synthesis and release of both follicle-stimulating hormone (FSH) and luteinizing hormone (LH) from the anterior pituitary glands. Emotional stress communicated to the hypothalamus can depress the formation and release of GnRH, preventing the release of FSH and LH. As a result, ovulation does not occur, progesterone is not produced, and amenorrhea results

 

Central Nervous System

Normal events and stimuli

Hypothalamus

GnRH

(anterior pituitary)

FSH

LH

Graafian follicle

Graafian follicle

Estrogen

prepare endometrium

Corpus luteum

Ovulation

Progesterone

(some estrogen)

 

prepare endometrium

 

Menstruation

 

 

 

 

 

 

 

Follicle-stimulating Hormone (FSH)

 

Female Ovary

 

Growth and maturation of graafian follicles and preparation of these cells so that they can respond to LH

 

luteinizing Hormone (LH)

 

Female Ovary

 

Works with FSH to stimulate estrogen secretion. Along with a surge of LH, estrogen stimulates ovulation, then luteinizes the follicle to form the corpus luteum produces progesterone and estrogen.

 

Hypothalamic-Pituitary Cycle

 

Toward the end of the normal menstrual cycle, blood levels of estrogen and progesterone fall. Low blood level of these ovarian hormones stimulate the hypothalamus to secrete GnRH, which in turn stimulates anterior pituitary secretion of FSH. FSH stimulates development of ovarian graafian follicles and their production of estrogen. Estrogen levels begin to fall and hypothalamic GnRH triggers the anterior pituitary release of LH. A marked surge of LH and smaller peak of estrogen precede the expulsion of the ovum from the graafian follicle by about 24 to 36 hours.

 

Ovarian Cycle

 

The primary graafian follicles contain immature oocytes. Before ovulation, from 1 to 30 follicles begin to mature in each ovary under the influence of FSH and estrogen. The preovulatory surge of LH affects a selected follicle. Within the chosen follicle, the oocyte matures, ovulation occurs, and the empty follicle begins its transformation into the corpus luteum (yellow body).

This follicular phase (preovulatory phase) of the ovarian menstrual cycle varies in length from woman to woman. Almost all variations in cycle length are the result of variations in the length of the follicular phase. On rare occasions (1 in 100 menstrual cycles), more than one follicle is chosen and more than one oocyte mature and undergoes ovulation (twins).

 

The events following ovulation, the luteal phase (postovulatory phase) of the ovarian menstrual cycle, usually require 14 days (range of 13 to 15 days). Eight days after ovulation, the corpus luteum reaches its peak of functional activity, secreting both of the steroids, estrogen and progesterone. Coincident with this time of peak luteal functioning, the fertilized egg is implanted in the endometrium. If no implantation occurs, the corpus luteum regresses, and steroid levels drop. Two weeks after ovulation, if fertilization and implantation do not occur, uterine endometrium is shed through menstruation.

 

After ovulation, estrogen levels drop. For 90% of women, only a small amount of withdrawal bleeding  occurs so that is goes unnoticed. In 10% of women, there is sufficient bleeding for it to be visible, resulting in what is known as midcycle bleeding

 

 

 

Endometrial Cycle

 

The endometrium responds to fluctuating levels of ovarian steroids. The endometrium consists of three layers. The basal layer intermingles with the myometrium. The other two functional layers undergo cyclic changes and are shed in menstrual flow.

 

When labeled according to phases in the endometrial cycle, the menstrual phase extends over the first 5 days when the two functional layers (the compact and spongy layers) are shed. The resting, or repair, phase ( the early proliferative phase) includes days 4 through 7.

The most rapid growth of the functional layers occurs between days 5 to 7 and the day of ovulation and is known as the proliferative phase. In a 28-day cycle, ovulation occurs on day 14 (range of 13 to 15 days). The secretory phase spans days 15 through 28; the last 3 days before menstruation make up the ischemic phase (stage of regression).

 

The endometrium has different characteristics at different times during the cycle depending on the levels of estrogen and progesterone. Therefore the endometrium can be “dated” (the phase of the cycle can be determined) and the presence and levels of hormones are too low or too high, the endometrium would reflect this.

 

The first day of menstrual discharge has been designated as day 1 of the cycle. The average duration of menstrual flow is 5 days (range of 3 to 6 days), and the average blood loss is approximately 50 ml (range of 20 to 80 ml), but there is great variation. During menstruation, the average daily loss of iron is 0.5 to 1mg. If the woman’s usual blood loss is over 80 ml, she will most likely need iron supplementation to prevent secondary anemia.

 

Uterine discharge includes mucus and epithelial cells in addition to blood.

 

Ovulation occurs about 14 days before the start of the next menstrual flow, for example, day 10 of a 24-day cycle, day 14 of a 28-day cycle, or day 18 of a 32-day cycle.

 

The endometrium surface is completely restored in approximately 4 days or slightly before the menstrual flow stops. From this point on, an eight fold to tenfold thickening occurs, with a leveling off of growth at ovulation. During the proliferative phase, the glands are tubular or columnar in shape. About 3 to 4 days before ovulation, vascularity to the endometrium increases. Endometrial growth during proliferation is dependent on estrogen produced by ovarian follicles before ovulation.

 

After ovulation, the graafian follicle develops into the corpus luteum. The corpus luteum produces estrogen and large amounts of progesterone. Progesterone causes the blood vessels in the endometrium to dilate and assume a spiral or corkscrew shape. Endometrium glands elongate, become more active, and secrete a glycogen containing fluid. The endometrium reaches the thickness of heavy, soft velvet and becomes luxuriant with blood and glandular secretions, a suitable protective and nutritive bed for a fertilized ovum. In the fully matured secretory endometrium, three strata are noted: the two functional or compact layer and an intermediate or spongy layer; and basal or inner, inactive layer.

If fertilization does not occur, about 3 days before menstruation begins, the corpus luteum degenerates with  

 

Vasospasm of the spiral arteries occurs. This vasoconstriction results in ischemia, necrosis, and sloughing off of the upper two layers of the endometrium. Near the end of the secretory phase, just before the start of menstrual flow, regeneration begins from the retained basal layer. Rebuilding the endometrium from the basal layer upward is responsible for its healing and rejuvenation without scar formation.